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During his doctoral research at the MRC Laboratory of Molecular Biology, Cambridge (1965-1969), he completed the sequence and crystallographic structure determination of the enzyme elastase. Following postdoctoral studies at the University of Bristol, Berkeley, and Harvard, during which he changed fields to cell biology, studying membrane structure, he took a lectureship at Imperial College in Improve your students' scientific skills and report writing with achievable experiments and simple structured guidance.

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